Indeed , the effect of the mutation in OE5 was similar to © 2008 The Authors Journal compilation © 2008 Blackwell Publishing Ltd , Molecular Microbiology , 71 , 146 -- 157NagC , GalR and GalS repression at galP 153 that in the weakly binding OE2 and OE3 sites ( a mutation in OE4 has not yet been tested ) .