Besides the mgtA gene , only 2 of phoPQ were directly regulated by the PhoPQ two-component system organisms .
Besides the mgtA gene , only 2 of phoPQ were directly regulated by the PhoPQ two-component system in .
All of the genes belong to the phoPQ regulon although the pmrHFIJKL genes are regulated by PhoPQ indirectly through the PmrAB signal transduction system .