As shown in Fig. 1 ( a , b ) , we found a sequence homologous to the RcsB-binding site present in other well-characterized RcsB-regulated genes , located 272 nt upstream from the previously described dps +1 transcription site ( Mouslim & Groisman , 2003 ; Yoo et al. , 2007 ) .
Taken together , our results confirm that RcsB promotes dps expression by directly binding to the promoter region of dps .
To determine which of the RcsB phosphorylation pathway is involved in the control of dps expression , levels of b-galactosidase : : lacZYA was determined in tolB P double rcsB mutants .
To determine which of the RcsB phosphorylation pathway is involved in the control of dps expression , levels of b-galactosidase : : lacZYA was determined in tolB rcsB , tolB PrcsDB .
Although Yoo et al. reported that dps expression is induced during the stationary-phase by the Fur regulator , our results demonstrated that this gene is also regulated by RcsB .